Family: Theridiidae Sundevall, 1833 [urn:lsid:nmbe.ch:spiderfam:0099]
Detail
Gen. Asagena Sundevall, 1833 [urn:lsid:nmbe.ch:spidergen:04190]
Detail
type Phalangium phaleratum Panzer, 1801; gender feminine
N.B.: removed from the synonymy of Steatoda Sundevall, 1833 by Wunderlich, 2008b: 198 (contra Levi, 1957b: 375); re-synonymization with Steatoda (Murphy & Roberts, 2015: viii) not sufficiently justified and not accepted here; Breitling, 2019b: 26 discusses Asagena might be a subgenus of Steatoda (based mainly on COI, see FAQ 7).
In synonymy:
Asagena amurica Strand, 1907 = Asagena phalerata (Panzer, 1801) (Breitling, 2015b: 478).
Asagena nesiotes (Chamberlin, 1924, T from Steatoda) = Asagena fulva (Keyserling, 1884) (Levi, 1957b: 391, sub Steatoda).
Asagena parvula (Banks, 1898, T from Steatoda) = Asagena fulva (Keyserling, 1884) (Levi, 1957b: 391, sub Steatoda).
Asagena phalerata corsica (Simon, 1873, T from Steatoda) = Asagena phalerata (Panzer, 1801) (Knoflach, 1996d: 382, sub Steatoda).
Asagena phalerata seraiensis (Nosek, 1905, T from Steatoda) = Asagena phalerata (Panzer, 1801) (Knoflach, 1996d: 382, sub Steatoda).
Asagena venusta (Marx, 1898, T from Steatoda) = Asagena fulva (Keyserling, 1884) (Levi, 1957b: 391, sub Steatoda).
Asagena amurica Strand, 1907 = Asagena phalerata (Panzer, 1801) (Breitling, 2015b: 478).
Asagena nesiotes (Chamberlin, 1924, T from Steatoda) = Asagena fulva (Keyserling, 1884) (Levi, 1957b: 391, sub Steatoda).
Asagena parvula (Banks, 1898, T from Steatoda) = Asagena fulva (Keyserling, 1884) (Levi, 1957b: 391, sub Steatoda).
Asagena phalerata corsica (Simon, 1873, T from Steatoda) = Asagena phalerata (Panzer, 1801) (Knoflach, 1996d: 382, sub Steatoda).
Asagena phalerata seraiensis (Nosek, 1905, T from Steatoda) = Asagena phalerata (Panzer, 1801) (Knoflach, 1996d: 382, sub Steatoda).
Asagena venusta (Marx, 1898, T from Steatoda) = Asagena fulva (Keyserling, 1884) (Levi, 1957b: 391, sub Steatoda).
Nomina dubia:
Asagena brasiliana Simon, 1864: 163, f. 61 (f, Brazil) -- Bonnet, 1955: 753.
Asagena dubia (Hentz, 1847: 461, pl. 24, f. 24, f, USA, originally in Herpyllus) -- Levi, 1957b: 400, sub Steatoda.
Asagena brasiliana Simon, 1864: 163, f. 61 (f, Brazil) -- Bonnet, 1955: 753.
Asagena dubia (Hentz, 1847: 461, pl. 24, f. 24, f, USA, originally in Herpyllus) -- Levi, 1957b: 400, sub Steatoda.
Asagena americana
Emerton, 1882 |
| Canada, USA, Mexico, China
[urn:lsid:nmbe.ch:spidersp:008021]
Asagena americana Emerton, 1882: 23, pl. 4, f. 6 (Dm).
Asagena americana Keyserling, 1886b: 2, pl. 11, f. 135 (mf).
Asagena americana Simon, 1894a: 574, f. 586.
Asagena americana Emerton, 1902: 122, f. 287 (mf).
Asagena americana Bishop, 1925: 66, f. 1 (m).
Asagena americana Kaston, 1948: 73, f. 25-29 (mf, removed from S of A. dubia).
Steatoda americana Levi, 1957b: 400, f. 66-69 (mf).
Steatoda phalerata Hu & Wu, 1989: 132, f. 103.1-2 (f; misidentified per Song, Zhu & Chen, 1999: 128).
Steatoda americana Knoflach, 1996d: 395, f. 58-64, 69, 74 (mf).
Steatoda americana Zhu, 1998: 338, f. 226A-E (mf).
Steatoda americana Song, Zhu & Chen, 1999: 128, f. 67C-D, K-L (mf).
Steatoda americana Paquin & Dupérré, 2003: 220, f. 2462-2464 (mf).
Steatoda americana Agnarsson, 2004: 607, f. 92D (m).
Steatoda americana Agnarsson, Coddington & Knoflach, 2007: 346, f. 17-19 (m).
Steatoda americana Agnarsson & Coddington, 2008: 53, f. 1C-D (m).
Asagena americana Wunderlich, 2008b: 199, f. 35 (m, T from Steatoda).
Asagena americana Álvarez-Padilla & Hormiga, 2011a: 849, f. 136B, 138E (mf).
Asagena americana Zhang, Peng & Zhang, 2022: 30, f. 13A-C (f).
Asagena americana Keyserling, 1886b: 2, pl. 11, f. 135 (mf).
Asagena americana Simon, 1894a: 574, f. 586.
Asagena americana Emerton, 1902: 122, f. 287 (mf).
Asagena americana Bishop, 1925: 66, f. 1 (m).
Asagena americana Kaston, 1948: 73, f. 25-29 (mf, removed from S of A. dubia).
Steatoda americana Levi, 1957b: 400, f. 66-69 (mf).
Steatoda phalerata Hu & Wu, 1989: 132, f. 103.1-2 (f; misidentified per Song, Zhu & Chen, 1999: 128).
Steatoda americana Knoflach, 1996d: 395, f. 58-64, 69, 74 (mf).
Steatoda americana Zhu, 1998: 338, f. 226A-E (mf).
Steatoda americana Song, Zhu & Chen, 1999: 128, f. 67C-D, K-L (mf).
Steatoda americana Paquin & Dupérré, 2003: 220, f. 2462-2464 (mf).
Steatoda americana Agnarsson, 2004: 607, f. 92D (m).
Steatoda americana Agnarsson, Coddington & Knoflach, 2007: 346, f. 17-19 (m).
Steatoda americana Agnarsson & Coddington, 2008: 53, f. 1C-D (m).
Asagena americana Wunderlich, 2008b: 199, f. 35 (m, T from Steatoda).
Asagena americana Álvarez-Padilla & Hormiga, 2011a: 849, f. 136B, 138E (mf).
Asagena americana Zhang, Peng & Zhang, 2022: 30, f. 13A-C (f).
Asagena brignolii
(Knoflach, 1996) |
| Greece
[urn:lsid:nmbe.ch:spidersp:008035]
Steatoda brignolii Knoflach, 1996d: 394, f. 54-57 (Df).
Asagena brignolii Wunderlich, 2008b: 199 (T from Steatoda).
Steatoda brignolii Le Peru, 2011: 464, f. 778 (f).
Asagena brignolii Wunderlich, 2008b: 199 (T from Steatoda).
Steatoda brignolii Le Peru, 2011: 464, f. 778 (f).
Asagena fulva
(Keyserling, 1884) |
| USA, Mexico
[urn:lsid:nmbe.ch:spidersp:008053]
Lithyphantes fulvus Keyserling, 1884a: 142, pl. 6, f. 89 (Df).
Lithyphantes parvula Banks, 1898b: 238, pl. 14, f. 1 (Dmf) [urn:lsid:nmbe.ch:spidersp:061566].
Lithyphantes venusta Marx, in Banks, 1898b: 239, pl. 14, f. 2 (Df) [urn:lsid:nmbe.ch:spidersp:061569].
Teutana nesiotes Chamberlin, 1924b: 639, f. 80 (Df) [urn:lsid:nmbe.ch:spidersp:061565].
Steatoda fulva Levi, 1957b: 391, f. 32-33, 45-47, 52 (mf, S of Steatoda nesiotes, S. parvula and S. venusta).
Steatoda fulva Gertsch, 1960b: 45, f. 62, 64-65, 70-71 (mf).
Asagena fulva Wunderlich, 2008b: 199 (T from Steatoda).
Asagena fulva Sánchez-Vega, Villegas-Guzmán & Durán-Barrón, 2019: 163, f. 1-3 (m).
Lithyphantes parvula Banks, 1898b: 238, pl. 14, f. 1 (Dmf) [urn:lsid:nmbe.ch:spidersp:061566].
Lithyphantes venusta Marx, in Banks, 1898b: 239, pl. 14, f. 2 (Df) [urn:lsid:nmbe.ch:spidersp:061569].
Teutana nesiotes Chamberlin, 1924b: 639, f. 80 (Df) [urn:lsid:nmbe.ch:spidersp:061565].
Steatoda fulva Levi, 1957b: 391, f. 32-33, 45-47, 52 (mf, S of Steatoda nesiotes, S. parvula and S. venusta).
Steatoda fulva Gertsch, 1960b: 45, f. 62, 64-65, 70-71 (mf).
Asagena fulva Wunderlich, 2008b: 199 (T from Steatoda).
Asagena fulva Sánchez-Vega, Villegas-Guzmán & Durán-Barrón, 2019: 163, f. 1-3 (m).
Asagena italica
(Knoflach, 1996) |
| France (incl. Corsica), Switzerland, Italy, Algeria
[urn:lsid:nmbe.ch:spidersp:008063]
Steatoda italica Knoflach, 1996d: 391, f. 3-4, 9, 11, 15-16, 18-20, 33-35, 39-41, 48-53, 65, 68, 73 (Dmf).
Asagena italica Wunderlich, 2008b: 199 (T from Steatoda).
Steatoda italica Le Peru, 2011: 465, f. 783, 791 (mf).
Asagena italica Iorio, 2023: 29, f. 22, 23a-b (m).
Asagena italica Wunderlich, 2008b: 199 (T from Steatoda).
Steatoda italica Le Peru, 2011: 465, f. 783, 791 (mf).
Asagena italica Iorio, 2023: 29, f. 22, 23a-b (m).
Asagena medialis
(Banks, 1898) |
| USA, Mexico
[urn:lsid:nmbe.ch:spidersp:008079]
Lithyphantes medialis Banks, 1898b: 240, pl. 14, f. 3 (Df).
Lithyphantes punctulatus Chamberlin, 1924b: 640 (S, rejected).
Steatoda medialis Levi, 1957b: 388, f. 42-43, 53-54 (f).
Steatoda medialis Gertsch, 1960b: 16, f. 6-10, 14-17, 28-30 (Dm, f).
Asagena medialis Wunderlich, 2008b: 199 (T from Steatoda).
Lithyphantes punctulatus Chamberlin, 1924b: 640 (S, rejected).
Steatoda medialis Levi, 1957b: 388, f. 42-43, 53-54 (f).
Steatoda medialis Gertsch, 1960b: 16, f. 6-10, 14-17, 28-30 (Dm, f).
Asagena medialis Wunderlich, 2008b: 199 (T from Steatoda).
Asagena meridionalis
Kulczyński, 1894 |
| Central to south-eastern and eastern Europe, Caucasus (Russia, Georgia)
[urn:lsid:nmbe.ch:spidersp:008080]
Asagena meridionalis Kulczyński, in Chyzer & Kulczyński, 1894: 39, pl. 2, f. 5 (Dm).
Asagena meridionalis Prisnij, 1981: 201, f. 1-9 (m, Df).
Steatoda meridionalis Brignoli, 1984b: 295, f. 17-18 (Df).
Steatoda meridionalis Knoflach, 1996d: 388, f. 2, 36-38, 42-47, 67, 72 (mf).
Steatoda meridionalis Svatoň, Prídavka & Pekár, 2002: 300, f. 1-3 (mf).
Asagena meridionalis Wunderlich, 2008b: 199 (T from Steatoda).
Steatoda meridionalis Le Peru, 2011: 466, f. 787, 791 (mf).
Asagena meridionalis Prisnij, 1981: 201, f. 1-9 (m, Df).
Steatoda meridionalis Brignoli, 1984b: 295, f. 17-18 (Df).
Steatoda meridionalis Knoflach, 1996d: 388, f. 2, 36-38, 42-47, 67, 72 (mf).
Steatoda meridionalis Svatoň, Prídavka & Pekár, 2002: 300, f. 1-3 (mf).
Asagena meridionalis Wunderlich, 2008b: 199 (T from Steatoda).
Steatoda meridionalis Le Peru, 2011: 466, f. 787, 791 (mf).
Asagena phalerata
(Panzer, 1801) *
|
| Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Iran, Central Asia, China, Korea
[urn:lsid:nmbe.ch:spidersp:008101]
Phalangium phaleratum Panzer, 1801: 78, pl. 21 (D).
Aranea signata Walckenaer, 1802: 209 (D).
Aranea serratipes Schrank, 1803: 233 (Dm).
Aranea notata Latreille, 1804a: 234 (D).
Theridion signatum Walckenaer, 1805: 76.
Drassus phalerata Sundevall, 1831: 26, 1832: 133.
Theridion 4-signatum Hahn, 1833a: 80, f. 60 (Dmf).
Asagena [phalerata] Sundevall, 1833b: 19.
Theridion quadrisignatum Hahn, 1836: 1, pl. 29, f. d (m).
Asagena serratipes C. L. Koch, 1839a: 98, f. 502-503 (m, Df).
Theridion signatum Walckenaer, 1841: 333 (mf).
Phrurolithus pallipes C. L. Koch, 1845: 98, f. 1026 (Df).
Latrodectus spinipes Lucas, 1846: 235, pl. 14, f. 9 (Dm).
Theridium serratipes Westring, 1861: 173.
Theridion signatum Blackwall, 1864a: 205, pl. 14, f. 135 (mf).
Asagena serratipes Menge, 1869: 256, pl. 49, f. 152 (mf).
Asagena corsica Simon, 1873a: 78 (Dmf) [urn:lsid:nmbe.ch:spidersp:061567].
Asagena phalerata Simon, 1881a: 173.
Asagena phalerata corsica Simon, 1881a: 174.
Asagena phalerata Hansen, 1882: 36, pl. 2, f. 13 (mf).
Asagena phalerata Chyzer & Kulczyński, 1894: 39, pl. 2, f. 4 (mf).
Asagena phalerata Becker, 1896: (2): 119, pl. 25, f. 4 (mf).
Asagena phalerata Bösenberg, 1902: 117, pl. 10, f. 148 (mf).
Asagena phalerata seraiensis Nosek, 1905b: 131 (Df) [urn:lsid:nmbe.ch:spidersp:061568].
Asagena amurica Strand, 1907k: 132 (Djf) [urn:lsid:nmbe.ch:spidersp:008022].
Asagena phalerata Reimoser, 1931b: 59, f. 3 (mf).
Asagena phalerata Wiehle, 1937: 202, f. 228-236 (mf).
Asagena phalerata Locket & Millidge, 1953: 54, f. 36A-C (mf).
Asagena phalerata Beer, 1964: 527, f. 2-3 (f).
Asagena phalerata Azheganova, 1968: 49, f. 83, 92 (mf).
Asagena phalerata Miller, 1971: 186, pl. XXXII, f. 4-5 (mf).
Steatoda phalerata Palmgren, 1974b: 40, f. 10.9-11, 18 (mf).
Asagena phalerata Xia & Zhu, 1980: 31, f. 1A-B (f).
Asagena phalerata Prisnij, 1981: 201, f. 10-18 (mf).
Asagena phalerata Hu, 1984: 158, f. 162.1-2 (f).
Steatoda phalerata Roberts, 1985: 178, f. 79c (mf).
Steatoda phalerata Zhang, 1987: 110, f. 90.1-2 (f).
Steatoda phalerata Hu & Wu, 1989: 132, f. 103.1-2 (f).
Steatoda phalerata Heimer & Nentwig, 1991: 296, f. 796 (mf).
Steatoda phalerata Roberts, 1995: 275, f. (mf).
Steatoda phalerata Namkung, Im & Kim, 1996: 15, f. 1-6 (m).
Steatoda phalerata Knoflach, 1996d: 382, f. 1, 5-8, 10, 12-14, 17, 21-32, 66, 70-71 (mf, S of Steatoda phalerata corsica and S. phalerata seraiensis).
Asagena phalerata Mcheidze, 1997: 183, f. 362-365 (mf).
Steatoda phalerata Zhu, 1998: 337, f. 225A-E (mf).
Steatoda phalerata Roberts, 1998: 290, f. (mf).
Steatoda phalerata Song, Zhu & Chen, 1999: 132, f. 69G-H, O-P (mf).
Steatoda phalerata Hu, 2001: 575, f. 388.1-2 (f).
Steatoda phalerata Song, Zhu & Chen, 2001: 107, f. 52A-E (mf).
Steatoda phalerata Namkung, 2002: 129, f. 13.47a-d (mf).
Steatoda phalerata Namkung, 2003: 131, f. 13.47a-d (mf).
Steatoda phalerata Almquist, 2006: 94, f. 116a-g (mf).
Steatoda phalerata Agnarsson, Coddington & Knoflach, 2007: 350, f. 39-42, 174-175 (m).
Asagena phalerata Wunderlich, 2008b: 199, f. 32-34 (m).
Steatoda phalerata Le Peru, 2011: 467, f. 790-791 (mf).
Asagena phalerata Kaya & Ugurtas, 2011: 146, f. 2-3 (mf).
Asagena phalerata Breitling, 2015b: 478 (S of Asagena amurica, removed from S of Steatoda albomaculata, rejecting Marusik, Mikhailov & Omelko, 2015: 123).
Asagena phalerata Vanuytven, 2021: 27, 77, f. A.13b, B.33b, B.34-35 (m).
Asagena phalerata Kim, 2021: 26, f. 7A-E, pl. 3 (m).
Aranea signata Walckenaer, 1802: 209 (D).
Aranea serratipes Schrank, 1803: 233 (Dm).
Aranea notata Latreille, 1804a: 234 (D).
Theridion signatum Walckenaer, 1805: 76.
Drassus phalerata Sundevall, 1831: 26, 1832: 133.
Theridion 4-signatum Hahn, 1833a: 80, f. 60 (Dmf).
Asagena [phalerata] Sundevall, 1833b: 19.
Theridion quadrisignatum Hahn, 1836: 1, pl. 29, f. d (m).
Asagena serratipes C. L. Koch, 1839a: 98, f. 502-503 (m, Df).
Theridion signatum Walckenaer, 1841: 333 (mf).
Phrurolithus pallipes C. L. Koch, 1845: 98, f. 1026 (Df).
Latrodectus spinipes Lucas, 1846: 235, pl. 14, f. 9 (Dm).
Theridium serratipes Westring, 1861: 173.
Theridion signatum Blackwall, 1864a: 205, pl. 14, f. 135 (mf).
Asagena serratipes Menge, 1869: 256, pl. 49, f. 152 (mf).
Asagena corsica Simon, 1873a: 78 (Dmf) [urn:lsid:nmbe.ch:spidersp:061567].
Asagena phalerata Simon, 1881a: 173.
Asagena phalerata corsica Simon, 1881a: 174.
Asagena phalerata Hansen, 1882: 36, pl. 2, f. 13 (mf).
Asagena phalerata Chyzer & Kulczyński, 1894: 39, pl. 2, f. 4 (mf).
Asagena phalerata Becker, 1896: (2): 119, pl. 25, f. 4 (mf).
Asagena phalerata Bösenberg, 1902: 117, pl. 10, f. 148 (mf).
Asagena phalerata seraiensis Nosek, 1905b: 131 (Df) [urn:lsid:nmbe.ch:spidersp:061568].
Asagena amurica Strand, 1907k: 132 (Djf) [urn:lsid:nmbe.ch:spidersp:008022].
Asagena phalerata Reimoser, 1931b: 59, f. 3 (mf).
Asagena phalerata Wiehle, 1937: 202, f. 228-236 (mf).
Asagena phalerata Locket & Millidge, 1953: 54, f. 36A-C (mf).
Asagena phalerata Beer, 1964: 527, f. 2-3 (f).
Asagena phalerata Azheganova, 1968: 49, f. 83, 92 (mf).
Asagena phalerata Miller, 1971: 186, pl. XXXII, f. 4-5 (mf).
Steatoda phalerata Palmgren, 1974b: 40, f. 10.9-11, 18 (mf).
Asagena phalerata Xia & Zhu, 1980: 31, f. 1A-B (f).
Asagena phalerata Prisnij, 1981: 201, f. 10-18 (mf).
Asagena phalerata Hu, 1984: 158, f. 162.1-2 (f).
Steatoda phalerata Roberts, 1985: 178, f. 79c (mf).
Steatoda phalerata Zhang, 1987: 110, f. 90.1-2 (f).
Steatoda phalerata Hu & Wu, 1989: 132, f. 103.1-2 (f).
Steatoda phalerata Heimer & Nentwig, 1991: 296, f. 796 (mf).
Steatoda phalerata Roberts, 1995: 275, f. (mf).
Steatoda phalerata Namkung, Im & Kim, 1996: 15, f. 1-6 (m).
Steatoda phalerata Knoflach, 1996d: 382, f. 1, 5-8, 10, 12-14, 17, 21-32, 66, 70-71 (mf, S of Steatoda phalerata corsica and S. phalerata seraiensis).
Asagena phalerata Mcheidze, 1997: 183, f. 362-365 (mf).
Steatoda phalerata Zhu, 1998: 337, f. 225A-E (mf).
Steatoda phalerata Roberts, 1998: 290, f. (mf).
Steatoda phalerata Song, Zhu & Chen, 1999: 132, f. 69G-H, O-P (mf).
Steatoda phalerata Hu, 2001: 575, f. 388.1-2 (f).
Steatoda phalerata Song, Zhu & Chen, 2001: 107, f. 52A-E (mf).
Steatoda phalerata Namkung, 2002: 129, f. 13.47a-d (mf).
Steatoda phalerata Namkung, 2003: 131, f. 13.47a-d (mf).
Steatoda phalerata Almquist, 2006: 94, f. 116a-g (mf).
Steatoda phalerata Agnarsson, Coddington & Knoflach, 2007: 350, f. 39-42, 174-175 (m).
Asagena phalerata Wunderlich, 2008b: 199, f. 32-34 (m).
Steatoda phalerata Le Peru, 2011: 467, f. 790-791 (mf).
Asagena phalerata Kaya & Ugurtas, 2011: 146, f. 2-3 (mf).
Asagena phalerata Breitling, 2015b: 478 (S of Asagena amurica, removed from S of Steatoda albomaculata, rejecting Marusik, Mikhailov & Omelko, 2015: 123).
Asagena phalerata Vanuytven, 2021: 27, 77, f. A.13b, B.33b, B.34-35 (m).
Asagena phalerata Kim, 2021: 26, f. 7A-E, pl. 3 (m).
Asagena pulchra
(Keyserling, 1884) |
| USA
[urn:lsid:nmbe.ch:spidersp:008104]
Lithyphantes pulcher Keyserling, 1884a: 137, pl. 6, f. 85 (Df).
Lithyphantes pulcher Banks, 1898b: 238, pl. 14, f. 7-8 (f).
Steatoda pulcher Levi, 1957b: 393, f. 37-38, 49 (f, Dm).
Steatoda pulcher Gertsch, 1960b: 33, f. 47-50, 55-56, 72-73 (mf).
Asagena pulcher Wunderlich, 2008b: 199 (T from Steatoda).
Lithyphantes pulcher Banks, 1898b: 238, pl. 14, f. 7-8 (f).
Steatoda pulcher Levi, 1957b: 393, f. 37-38, 49 (f, Dm).
Steatoda pulcher Gertsch, 1960b: 33, f. 47-50, 55-56, 72-73 (mf).
Asagena pulcher Wunderlich, 2008b: 199 (T from Steatoda).
Asagena semideserta
(Ponomarev, 2005) |
| Kazakhstan, Mongolia
[urn:lsid:nmbe.ch:spidersp:039813]
Steatoda semideserta Ponomarev, 2005: 47, f. 4d-g (Dmf).
Asagena semideserta Marusik, Fomichev & Tuneva, 2016: 478, f. 1-28 (mf, T from Steatoda).
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Asagena semideserta Marusik, Fomichev & Tuneva, 2016: 478, f. 1-28 (mf, T from Steatoda).